A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order Anura (coming from the Ancient Greek ἀνούρα, literally 'without tail'). Frog species with rough skin texture due to wart-like parotoid glands tend to be called toads, but the distinction between frogs and toads is informal and purely cosmetic, and does not reflect taxonomy or evolutionary history. Frogs are widely distributed, ranging from the tropics to subarctic regions, but the greatest concentration of species diversity is in tropical rainforest and associated wetlands. They account for around 88% of extant amphibian species, and are one of the five most diverse vertebrate orders. The oldest fossil "proto-frog" Triadobatrachus is known from the Early Triassic of Madagascar (250 million years ago), but molecular clock dating suggests their divergence from other amphibians may extend further back to the Permian, 265 million years ago. Adult frogs have a stout body, protruding eyes, anteriorly-attached tongue, limbs folded underneath, and no tail (the "tail" of tailed frogs is an extension of the male cloaca). Frogs have glandular skin, with secretions ranging from distasteful to toxic. Their skin varies in colour from well-camouflaged dappled brown, grey and green, to vivid patterns of bright red or yellow and black to show toxicity and ward off predators. Adult frogs live in both fresh water and on dry land; some species are adapted for living underground or in trees. As their skin is semi-permeable, making them susceptible to dehydration, they either live in moist niches or have special adaptations to deal with drier habitats.